Devil May Cry 4: Special Edition follows the story of Nero, a young knight of the Order of the Sword who possesses a mysterious demonic arm known as the Devil Bringer. During a sacred ceremony, the legendary demon hunter Dante suddenly attacks the Order and kills its leader, forcing Nero to pursue him and uncover the truth behind his actions. As the journey unfolds, Nero discovers a dangerous conspiracy involving the exploitation of demonic power and the Order's hidden ambitions. The story later shifts perspectives, allowing players to experience events through Dante and additional playable characters, each with their own unique abilities and motivations. Along the way, powerful enemies and ancient demons threaten the balance between the human and demon worlds. Themes of destiny, family, and the struggle between humanity and demonic heritage are central to the narrative. The Special Edition expands the original experience by introducing new playable characters and deeper insight into the events surrounding Fortuna and its inhabitants.